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In ferns, applied to the supposedly primitive, massive, stout-stalked, thick-walled type of sporangium. This arises from more than one parent cell, and lacks the specialized annulus or ring of thickened cells that is present in the leptosporangiate, supposedly more advanced ferns. (The latter have delicate, thin-walled, and thin-stalked sporangia.)

Subjects: Plant Sciences and Forestry.

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