1 An age (40.4–48.6 Ma ago) in the Middle Eocene, preceded by the Ypresian (Early Eocene), followed by the Bartonian (Int. Commission on Stratigraphy, 2004).
2 The name of the corresponding European stage, which is roughly contemporaneous with the upper Ulatisian and lower Narizian (N. America), the upper Heretaungan, Porangan, and lower Bortonian (New Zealand), and the upper Johannian and lower Aldingan (Australia).
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