Northern Plains tribe affiliated with the Arikara, combined bison hunting with agricultural pursuits. La Vérendrye visited the Mandan in 1738, and every later expedition up the Missouri knew them. By 1776 smallpox and attacks by neighboring tribes had reduced them to a single village. In 1870 they settled with other tribes on the Fort Berthold reservation in North Dakota. The Mandan figure in the journals of Lewis and Clark and are the subject of a study and many paintings by George Catlin.