1 The earliest age in the Eocene Epoch, preceded by the Thanetian (Palaeocene), followed by the Lutetian, and dated at 55.8–48.6 Ma (Int. Commission on Stratigraphy, 2004).
2 The name of the corresponding European stage, named after the Ypres Clay, Belgium. It is roughly contemporaneous with the Penutian and lower Ulatisian (N. America), upper Waipawan, Mangaorapan and lower Heretaungan (New Zealand), and the lower Johannian (Australia).
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