Chapter

Phylogeny of Heliodinidae

Yu-Feng Hsu and Jerry A. Powell

in Phylogenetic Relationships within Heliodinidae and Systematics of Moths Formerly Assigned to Heliodines Stainton (Lepidoptera: Yponomeutoidea)

Published by University of California Press

Published in print January 2005 | ISBN: 9780520098473
Published online March 2012 | e-ISBN: 9780520916067 | DOI: http://dx.doi.org/10.1525/california/9780520098473.003.0004
Phylogeny of Heliodinidae

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Systematists have relied largely on forewing venation, color and scaling on forewing and legs, and perching behavior to characterize heliodinids. These features, however, are not unique to the family and have been misinterpreted. Consequently, members of non-heliodinid moths have been placed in the family, while certain heliodinids have been placed in other families such as Glyphipterigidae. Although these features enable confident recognition of most members of the family, they are not appropriate for cladistic definition of Heliodinidae. Kyrki (1984, 1990) proposed a phylogenetic hypothesis of the superfamily Yponomeutoidea. Four characters were listed by Kyrki (1984) as possible autapomorphies for Heliodinidae: smooth scaling on the head; vein CuP absent from the forewing; strong lateral ridges on the pupa; and stiff, long lateral and dorsal bristles on the pupa. Two of Kyrki's (1990) four characters (the first and last) plus the three additional autapomorphies of Heliodinidae proposed in this chapter were included as characters 32 through 36 in the data matrix for the cladistic analysis to affirm the monophyly of Heliodinidae from the yponomeutoid outgroups.

Keywords: Heliodinidae; heliodinids; monophyly; cladistic analysis; moths; Yponomeutoidea; autapomorphies; pupa

Chapter.  5359 words.  Illustrated.

Subjects: Zoology and Animal Sciences

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